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Prior to the events of the novel, Moslov was a star pupil at the basic training school near Novosibirsk, and worked for some time at the Technical Support Group of the KGB’s Second Chief Directorate.
Following a deadly encounter with Russian knifemen, M assigns James Bond on a covert operation codenamed Operation Icebreaker in cooperation with agents of the KGB, CIA, and Mossad. Representing the KGB, Koyla Mosolov tells the three agents that they are pursuing the National Socialist Action Army for the theft of arms stolen from Russian bases, and to investigate the future destination of the stolen arms.
Following Rivke Ingber's accident in a land mine explosion, Koyla shows his CIA and MI6 counterparts a map where he has von Glöda's headquarters targeted in Blue Hare near Alakurtti within the Soviet Union. When they arrive at the Ice Palace, Bond learns from the self-appointed Führer that he collaborated with Mosolov revealing that in exchange for his service to him, he would give Bond over to him so he can fall into the custody of Department V (formerly known as SMERSH). After being tortured for valuable information, Bond escapes with Paula Vacker to her headquarters where Mosolov finds them to alert them that the Soviet Union has launched airstrikes at the Ice Palace demolishing it in the process. However, von Glöda escapes.
Now back in Helsinki, Bond questions Vacker about her true affiliations where Vacker discloses that she worked under orders of Mosolov and SUPO, and the knife incident occurred when SUPO informed Mosolov of Bond's arrival. That night, while Bond is sleeping with Vacker, he is confronted by Koslov again at gunpoint who reveals he had administrated Vacker with a small injection that will leave her unconscious for six to seven hours. He also reveals von Glöda is leaving Finland through the Vantaan Airport en route to London to stage a rally. Then, he orders Bond to his Saab where he hopes to use his built-in telephone to alert von Glöda of Bond's capture. There, Bond kills the KGB agent with his magnum cartridge, and drives to the airport to confront von Glöda.